:root {
    --canvas: #f7f7f4;
    --canvas-soft: #fafaf7;
    --surface-card: #ffffff;
    --surface-strong: #e6e5e0;
    --primary: #f54e00;
    --primary-active: #d04200;
    --on-primary: #ffffff;
    --ink: #26251e;
    --body: #5a5852;
    --body-strong: #26251e;
    --muted: #807d72;
    --muted-soft: #a09c92;
    --hairline: #e6e5e0;
    --hairline-soft: #efeee8;
    --hairline-strong: #cfcdc4;
    --semantic-success: #1f8a65;
    --semantic-error: #cf2d56;
    --rounded-xs: 4px;
    --rounded-sm: 6px;
    --rounded-md: 8px;
    --rounded-lg: 12px;
    --rounded-xl: 16px;
    --rounded-pill: 9999px;
    --spacing-xxs: 4px;
    --spacing-xs: 8px;
    --spacing-sm: 12px;
    --spacing-base: 16px;
    --spacing-md: 20px;
    --spacing-lg: 24px;
    --spacing-xl: 32px;
    --spacing-xxl: 48px;
    --spacing-section: 80px;
    --font-sans: 'Inter', system-ui, 'Helvetica Neue', Helvetica, Arial, sans-serif;
    --font-mono: 'JetBrains Mono', 'Courier New', monospace;
}

*, *::before, *::after { box-sizing: border-box; margin: 0; padding: 0; }

html { font-size: 16px; scroll-behavior: smooth; }

body {
    background: var(--canvas);
    color: var(--body);
    font-family: var(--font-sans);
    font-size: 16px;
    font-weight: 400;
    line-height: 1.5;
    -webkit-font-smoothing: antialiased;
}

.container {
    max-width: 1200px;
    margin: 0 auto;
    padding: 0 var(--spacing-xl);
}

img { display: block; max-width: 100%; height: auto; }

a { color: var(--ink); text-decoration: underline; text-underline-offset: 2px; }
a:hover { color: var(--primary); }

h1, h2, h3, h4 {
    color: var(--ink);
    font-weight: 400;
    line-height: 1.2;
    letter-spacing: -0.02em;
}
h1 { font-size: 56px; letter-spacing: -0.03em; line-height: 1.1; }
h2 { font-size: 36px; letter-spacing: -0.02em; margin-bottom: var(--spacing-lg); }
h3 { font-size: 22px; letter-spacing: -0.01em; margin-bottom: var(--spacing-sm); }

p { margin-bottom: var(--spacing-base); }
p:last-child { margin-bottom: 0; }

ul, ol { padding-left: var(--spacing-lg); }
li { margin-bottom: var(--spacing-xs); }

address { font-style: normal; }
address p { margin-bottom: var(--spacing-xs); }

em { font-style: italic; }
strong { font-weight: 600; color: var(--body-strong); }

code {
    font-family: var(--font-mono);
    font-size: 13px;
    background: var(--surface-card);
    border: 1px solid var(--hairline);
    border-radius: var(--rounded-xs);
    padding: 1px 5px;
    color: var(--ink);
}

.site-header {
    position: sticky;
    top: 0;
    z-index: 100;
    background: var(--canvas);
    border-bottom: 1px solid var(--hairline);
    height: 64px;
}

.header-inner {
    display: flex;
    align-items: center;
    justify-content: space-between;
    height: 64px;
}

.site-logo {
    font-size: 18px;
    font-weight: 600;
    color: var(--ink);
    text-decoration: none;
    letter-spacing: -0.01em;
    flex-shrink: 0;
}
.site-logo:hover { color: var(--primary); }

.site-nav ul {
    display: flex;
    list-style: none;
    gap: var(--spacing-xl);
    padding: 0;
    margin: 0;
}

.site-nav a {
    font-size: 14px;
    font-weight: 500;
    color: var(--body);
    text-decoration: none;
    line-height: 1.4;
    transition: color 0.15s;
}
.site-nav a:hover,
.site-nav a[aria-current="page"] { color: var(--ink); }

.nav-toggle {
    display: none;
    flex-direction: column;
    justify-content: space-between;
    width: 24px;
    height: 18px;
    background: none;
    border: none;
    cursor: pointer;
    padding: 0;
}
.nav-toggle span {
    display: block;
    width: 100%;
    height: 2px;
    background: var(--ink);
    border-radius: 2px;
    transition: transform 0.2s, opacity 0.2s;
}

.badge-pill {
    display: inline-block;
    background: var(--surface-strong);
    color: var(--ink);
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    border-radius: var(--rounded-pill);
    padding: 4px 10px;
    margin-bottom: var(--spacing-lg);
}

.hero-band {
    padding: var(--spacing-section) 0 0;
}

.hero-band .container { max-width: 800px; }

.hero-band h1 {
    margin-bottom: var(--spacing-lg);
}

.hero-sub {
    font-size: 18px;
    color: var(--muted);
    line-height: 1.5;
    max-width: 640px;
    margin-bottom: var(--spacing-xl);
}

.hero-image-wrap {
    margin-top: var(--spacing-xl);
    border-top: 1px solid var(--hairline);
}

.hero-img {
    width: 100%;
    max-height: 480px;
    object-fit: cover;
    display: block;
}

.img-caption {
    font-size: 13px;
    color: var(--muted);
    padding: var(--spacing-sm) var(--spacing-xl);
    border-bottom: 1px solid var(--hairline);
}

.page-hero {
    padding: var(--spacing-xxl) 0;
    border-bottom: 1px solid var(--hairline);
}

.page-hero h1 { font-size: 40px; }

.section-band {
    padding: var(--spacing-section) 0;
    border-bottom: 1px solid var(--hairline-soft);
}

.section-band:last-of-type { border-bottom: none; }

.section-alt { background: var(--canvas-soft); }

.section-label {
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    color: var(--muted);
    margin-bottom: var(--spacing-sm);
}

.two-col-text {
    display: grid;
    grid-template-columns: 1fr 1fr;
    gap: var(--spacing-xl);
}

.two-col-text p { color: var(--body); }

.card-grid {
    display: grid;
    grid-template-columns: repeat(3, 1fr);
    gap: var(--spacing-lg);
    margin-top: var(--spacing-xl);
}

.feature-card {
    background: var(--surface-card);
    border: 1px solid var(--hairline);
    border-radius: var(--rounded-lg);
    overflow: hidden;
    display: flex;
    flex-direction: column;
}

.card-img-link { display: block; overflow: hidden; }
.card-img-link img {
    width: 100%;
    height: 220px;
    object-fit: cover;
    transition: transform 0.3s ease;
}
.feature-card:hover .card-img-link img { transform: scale(1.03); }

.card-body {
    padding: var(--spacing-lg);
    flex: 1;
    display: flex;
    flex-direction: column;
}

.card-label {
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    color: var(--muted);
    margin-bottom: var(--spacing-xs);
}

.card-body h3 {
    font-size: 18px;
    margin-bottom: var(--spacing-sm);
    flex: 1;
}

.card-body h3 a {
    color: var(--ink);
    text-decoration: none;
}
.card-body h3 a:hover { color: var(--primary); }

.card-body p {
    font-size: 14px;
    color: var(--body);
    margin-bottom: var(--spacing-base);
}

.card-link {
    font-size: 14px;
    font-weight: 500;
    color: var(--primary);
    text-decoration: none;
    display: inline-flex;
    align-items: center;
    gap: 4px;
    margin-top: auto;
}
.card-link::after { content: '→'; }
.card-link:hover { color: var(--primary-active); }

.species-table-wrap {
    overflow-x: auto;
    margin-top: var(--spacing-lg);
    border: 1px solid var(--hairline);
    border-radius: var(--rounded-lg);
}

.species-table {
    width: 100%;
    border-collapse: collapse;
    background: var(--surface-card);
    font-size: 14px;
}

.species-table th {
    text-align: left;
    padding: var(--spacing-sm) var(--spacing-base);
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    color: var(--muted);
    border-bottom: 1px solid var(--hairline);
    background: var(--canvas-soft);
}

.species-table td {
    padding: var(--spacing-sm) var(--spacing-base);
    border-bottom: 1px solid var(--hairline-soft);
    color: var(--body);
    vertical-align: top;
}

.species-table tr:last-child td { border-bottom: none; }
.species-table tbody tr:hover td { background: var(--canvas-soft); }

.table-note {
    font-size: 13px;
    color: var(--muted);
    margin-top: var(--spacing-sm);
}

.contact-section { max-width: 720px; }
.contact-intro { color: var(--muted); margin-bottom: var(--spacing-xl); }

.contact-form { display: flex; flex-direction: column; gap: var(--spacing-base); }

.form-group { display: flex; flex-direction: column; gap: var(--spacing-xs); }

.form-group label {
    font-size: 14px;
    font-weight: 500;
    color: var(--ink);
}

.form-group input,
.form-group textarea {
    background: var(--surface-card);
    border: 1px solid var(--hairline-strong);
    border-radius: var(--rounded-md);
    padding: 12px 16px;
    font-family: var(--font-sans);
    font-size: 16px;
    color: var(--ink);
    height: 44px;
    transition: border-color 0.15s;
    width: 100%;
}

.form-group textarea {
    height: auto;
    resize: vertical;
    min-height: 120px;
}

.form-group input:focus,
.form-group textarea:focus {
    outline: none;
    border-color: var(--ink);
}

.form-group input::placeholder,
.form-group textarea::placeholder { color: var(--muted-soft); }

.btn-primary {
    display: inline-block;
    background: var(--primary);
    color: var(--on-primary);
    font-family: var(--font-sans);
    font-size: 14px;
    font-weight: 500;
    padding: 10px 18px;
    height: 40px;
    border: none;
    border-radius: var(--rounded-md);
    cursor: pointer;
    text-decoration: none;
    line-height: 1;
    transition: background 0.15s;
    align-self: flex-start;
}
.btn-primary:hover { background: var(--primary-active); color: var(--on-primary); }

.btn-secondary {
    display: inline-block;
    background: var(--surface-card);
    color: var(--ink);
    font-family: var(--font-sans);
    font-size: 14px;
    font-weight: 500;
    padding: 10px 18px;
    height: 40px;
    border: 1px solid var(--hairline-strong);
    border-radius: var(--rounded-md);
    cursor: pointer;
    text-decoration: none;
    line-height: 1;
    transition: background 0.15s;
}
.btn-secondary:hover { background: var(--canvas-soft); color: var(--ink); }

.form-status {
    font-size: 14px;
    padding: var(--spacing-sm) 0;
    min-height: 24px;
}
.form-status.success { color: var(--semantic-success); }
.form-status.error { color: var(--semantic-error); }

.article-layout {
    display: grid;
    grid-template-columns: 1fr 320px;
    gap: var(--spacing-xxl);
    align-items: start;
}

.article-body { max-width: 720px; }

.article-lead {
    font-size: 18px;
    color: var(--body-strong);
    line-height: 1.6;
    margin-bottom: var(--spacing-xl);
    border-left: 3px solid var(--primary);
    padding-left: var(--spacing-base);
}

.article-body h2 {
    font-size: 28px;
    margin-top: var(--spacing-xxl);
    margin-bottom: var(--spacing-base);
    padding-top: var(--spacing-lg);
    border-top: 1px solid var(--hairline-soft);
}

.article-body h2:first-of-type { border-top: none; margin-top: var(--spacing-lg); }

.article-body h3 { font-size: 20px; margin-top: var(--spacing-xl); }

.article-body p { color: var(--body); }

.article-figure {
    margin: var(--spacing-xl) 0;
    border: 1px solid var(--hairline);
    border-radius: var(--rounded-lg);
    overflow: hidden;
    background: var(--surface-card);
}

.article-figure img {
    width: 100%;
    max-height: 420px;
    object-fit: cover;
}

.article-figure figcaption {
    font-size: 13px;
    color: var(--muted);
    padding: var(--spacing-sm) var(--spacing-base);
    border-top: 1px solid var(--hairline-soft);
    line-height: 1.4;
}

.content-list {
    color: var(--body);
    margin-bottom: var(--spacing-base);
}
.content-list li { margin-bottom: var(--spacing-sm); }

.info-block {
    background: var(--surface-card);
    border: 1px solid var(--hairline);
    border-left: 3px solid var(--primary);
    border-radius: var(--rounded-md);
    padding: var(--spacing-lg);
    margin: var(--spacing-xl) 0;
}

.info-block-label {
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    color: var(--muted);
    margin-bottom: var(--spacing-sm);
}

.info-block ol {
    margin: 0;
    color: var(--body);
}

.article-nav {
    margin-top: var(--spacing-xxl);
    padding-top: var(--spacing-lg);
    border-top: 1px solid var(--hairline);
}

.article-sidebar { position: sticky; top: 88px; }

.sidebar-card {
    background: var(--surface-card);
    border: 1px solid var(--hairline);
    border-radius: var(--rounded-lg);
    padding: var(--spacing-lg);
    margin-bottom: var(--spacing-base);
}

.sidebar-label {
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    color: var(--muted);
    margin-bottom: var(--spacing-sm);
}

.sidebar-card ul {
    list-style: none;
    padding: 0;
    margin: 0;
}

.sidebar-card li { margin-bottom: var(--spacing-sm); }

.sidebar-card a {
    font-size: 14px;
    color: var(--body);
    text-decoration: none;
    line-height: 1.4;
}
.sidebar-card a:hover { color: var(--primary); }

.content-narrow { max-width: 720px; }

.content-narrow h2 {
    margin-top: var(--spacing-xxl);
    padding-top: var(--spacing-lg);
    border-top: 1px solid var(--hairline-soft);
}
.content-narrow h2:first-of-type { border-top: none; margin-top: 0; }

.content-narrow .content-list {
    margin-bottom: var(--spacing-lg);
}

.site-footer {
    background: var(--canvas);
    border-top: 1px solid var(--hairline);
    padding: 64px 0 0;
}

.footer-inner {
    display: grid;
    grid-template-columns: 2fr 1fr 1fr 1fr 1fr;
    gap: var(--spacing-xxl);
    padding-bottom: var(--spacing-xxl);
}

.footer-brand {
    font-size: 16px;
    font-weight: 600;
    color: var(--ink);
    margin-bottom: var(--spacing-xs);
}

.footer-tagline {
    font-size: 14px;
    color: var(--muted);
    line-height: 1.4;
}

.footer-heading {
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    color: var(--muted);
    margin-bottom: var(--spacing-sm);
}

.footer-col ul {
    list-style: none;
    padding: 0;
    margin: 0;
}

.footer-col li { margin-bottom: var(--spacing-xs); }

.footer-col a {
    font-size: 14px;
    color: var(--body);
    text-decoration: none;
}
.footer-col a:hover { color: var(--ink); }

.footer-bottom {
    border-top: 1px solid var(--hairline);
    padding: var(--spacing-lg) 0;
}

.footer-bottom .container {
    display: flex;
    justify-content: space-between;
    align-items: center;
    flex-wrap: wrap;
    gap: var(--spacing-sm);
}

.footer-bottom p { font-size: 13px; color: var(--muted); margin: 0; }
.footer-bottom a { color: var(--body); text-decoration: none; }
.footer-bottom a:hover { color: var(--ink); }

.cookie-banner {
    position: fixed;
    bottom: var(--spacing-lg);
    left: 50%;
    transform: translateX(-50%);
    background: var(--ink);
    color: var(--canvas);
    border-radius: var(--rounded-lg);
    padding: var(--spacing-base) var(--spacing-lg);
    display: flex;
    align-items: center;
    gap: var(--spacing-lg);
    max-width: 680px;
    width: calc(100% - var(--spacing-xl));
    z-index: 999;
    box-shadow: 0 4px 24px rgba(38, 37, 30, 0.16);
    font-size: 14px;
}

.cookie-banner p { margin: 0; color: var(--canvas); }
.cookie-banner a { color: var(--canvas); }

.cookie-actions {
    display: flex;
    gap: var(--spacing-sm);
    flex-shrink: 0;
}

.cookie-banner .btn-primary {
    background: var(--canvas);
    color: var(--ink);
    font-size: 13px;
    height: 36px;
    padding: 8px 14px;
}
.cookie-banner .btn-primary:hover { background: var(--canvas-soft); }

.cookie-banner .btn-secondary {
    background: transparent;
    color: var(--canvas);
    border-color: rgba(255,255,255,0.3);
    font-size: 13px;
    height: 36px;
    padding: 8px 14px;
}
.cookie-banner .btn-secondary:hover { background: rgba(255,255,255,0.1); }

.cookie-hidden { display: none !important; }

@media (max-width: 1024px) {
    h1 { font-size: 40px; }
    h2 { font-size: 28px; }

    .two-col-text { grid-template-columns: 1fr; gap: var(--spacing-base); }
    .card-grid { grid-template-columns: repeat(2, 1fr); }
    .footer-inner { grid-template-columns: 1fr 1fr; gap: var(--spacing-xl); }
    .article-layout { grid-template-columns: 1fr; }
    .article-sidebar { position: static; }
}

@media (max-width: 768px) {
    .container { padding: 0 var(--spacing-base); }

    h1 { font-size: 32px; }
    h2 { font-size: 24px; }

    .hero-band { padding: var(--spacing-xxl) 0 0; }
    .section-band { padding: var(--spacing-xxl) 0; }
    .card-grid { grid-template-columns: 1fr; }
    .footer-inner { grid-template-columns: 1fr; gap: var(--spacing-lg); }

    .nav-toggle { display: flex; }
    .site-nav {
        display: none;
        position: absolute;
        top: 64px;
        left: 0;
        right: 0;
        background: var(--canvas);
        border-bottom: 1px solid var(--hairline);
        padding: var(--spacing-lg);
    }
    .site-nav.open { display: block; }
    .site-nav ul { flex-direction: column; gap: var(--spacing-base); }

    .footer-bottom .container { flex-direction: column; align-items: flex-start; }
    .species-table-wrap { border-radius: 0; border-left: none; border-right: none; }
}

@media (max-width: 480px) {
    h1 { font-size: 26px; }
    .cookie-banner { flex-direction: column; align-items: flex-start; gap: var(--spacing-sm); }
}